This is also the optimal time frame for turf inoculation or applications with milky spore to increase milky spore in the soil environment (there are product specific guidelines that should be followed for milky spore application). In August, when the grubs are close to the surface and feeding, they are vulnerable to infestation by milky spore.
As the weather gets cooler and winter approaches, the grubs go deeper into the soil, and feeding declines as they over-winter. The eggs hatch soon afterwards and in this larval or grub stage, they feed on the roots of grass and other plants. During this adult stage, the beetles also mate and the females lay eggs in the soil in late July to early August. The adult Japanese beetles pupate in July (in the Northeast United States) and feed on flowers and leaves of shrubs and garden plants. It is responsible for a disease (commonly called milky spore) of the white grubs of Japanese beetles. Paenibacillus popilliae (formerly Bacillus popilliae) is a soil-dwelling, Gram-positive, rod-shaped bacterium.
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